Mathematics

Resource-mediated competition between two plant species with different rates of water intake

Speaker: 
Chunyi Gai
Date: 
Tue, May 11, 2021
Location: 
UBC
Online
Conference: 
PIMS Workshop on New Trends in Localized Patterns in PDES
Abstract: 

We propose an extension of the well-known Klausmeier model of vegetation to two plant species that consume water at different rates. Rather than competing directly, the plants compete through their intake of water, which is a shared resource between them. In semi-arid regions, the Klausmeier model produces vegetation spot patterns. We are interested in how the competition for water affects co-existence and stability of patches of different plant species. We consider two plant types: a thirsty species and a frugal species, that only differ by the amount of water they consume, while being identical in all other aspects. We find that there is a finite range of precipitation rate for which two species can co-exist. Outside of that range, (when the rate is either sufficiently low or high), the frugal species outcompetes the thirsty species. As the precipitation rate is decreased, there is sequence of stability thresholds such that thirsty plant patches are the first to die off, while the frugal spots remain resilient for longer. The pattern consisting of only frugal spots is the most resilient. The next-most-resilient pattern consists of all-thirsty patches, with the mixed pattern being less resilient than either of the homogeneous patterns. We also examine numerically what happens for very large precipitation rate. We find that for sufficiently high rate, the frugal plant takes over the entire range, outcompeting the thirsty plant.

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Boundary Layer Solutions in the Gierer-Meinhardt System

Speaker: 
Daniel Gomez
Date: 
Tue, May 11, 2021
Location: 
UBC
Online
Conference: 
PIMS Workshop on New Trends in Localized Patterns in PDES
Abstract: 

The singularly perturbed Gierer-Meinhardt system has been a prototypical reaction diffusion system for the analysis of localized multi spike solutions. Motivated by recent interest in bulk-surface coupled systems, in this talk we address the structure and linear stability of multi spike solutions in the presence of inhomogeneous boundary conditions. Such inhomogeneities are shown to lead to the formation of both stable symmetric and asymmetric boundary bound spike solutions in one-dimensional domains and analogous solutions in higher dimensions.

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Noise and Spike Dynamics for the Gierer-Meinhardt Equations

Speaker: 
David Iron
Date: 
Mon, May 10, 2021
Location: 
UBC
Online
Conference: 
PIMS Workshop on New Trends in Localized Patterns in PDES
Abstract: 

The stability and dynamic properties of spike-type solutions to the Gierer- Meinhart equations are well understood. We examine the effect of adding noise to the equations on the spike-dynamics. We derive a stochastic ordinary differential equation for the motion of a single spike as well as the distribution of spike location from the associated Fokker-Plank equation. With sufficiently large amplitude noise, it is possible for the spike to reach the boundary of the domain and become effectively trapped for some time. In this case, we calculate the expected time to capture.

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A ring of spikes for the Schnakenberg model

Speaker: 
Theodore Kolokolnikov
Date: 
Mon, May 10, 2021
Location: 
UBC
Online
Conference: 
PIMS Workshop on New Trends in Localized Patterns in PDES
Abstract: 

Consider N spikes on located along a ring inside a unit disk. This highly symmetric configuration corresponds to an equilibrium of a two-dimensional Schnakenberg model; the ring radius can be characterized in terms of the modified Green’s function. We study the stability of such a ring with respect to both small and large eigenvalues (corresponding to spike position and spike height perturbations, respectively), and characterize the instability thresholds. For sufficiently large feed rate, we find that a ring of 8 or less spikes is stable with respect to both small and large eigenvalues, whereas a ring of 9 spikes is unstable with respect to small eigenvalues. For 8 spikes or less, as the feed rate is decreased, a small eigen-value instability is triggered first, followed by large eigenvalue instability. For 8 spikes, this instability appears to be supercritical, and deforms a ring into a square-type configuration. The main tool we use is circulant matrices and an analogue of the floquet theory.

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The Phase-Field-Crystal Model at Large and Small Scales

Speaker: 
Rustum Choksi
Date: 
Mon, May 10, 2021
Location: 
UBC
Online
Conference: 
PIMS Workshop on New Trends in Localized Patterns in PDES
Abstract: 

The Phase-Field-Crystal (PFC) model is a simple yet surprisingly useful model for successfully capturing the phenomenology of grain growth in polycrystalline materials. PFC models are variational with a free energy functional which is very similar (in some cases, identical) to the well-known Swift-Hohenberg free energy. In this talk, we will discuss the simplest PFC functional and its gradient flow.

The first part of the talk will focus on large scales and address the model’s uncanny ability to o capture certain features of grain growth. We introduce a novel atom-based grain extraction and measurement method, and then use it to provide a comparison of multiple statistical grain metrics between (i) PFC simulations, (ii) experimental thin film data for aluminum, and (iii) simulations from the Mullins model.

In the second part of the talk, we investigate the PFC energy landscape at small scales (the local arrangement of atoms). We address patterns which are numerically observed as steady states via the framework of the modern theory of rigorous computations. In doing so, we make rigorous conclusions on the existence of similar states. In particular, we show that localized patterns and grain boundaries are critical and not simply metastable states. Finally, we present preliminary work on connections and parameter continuation in the PFC system. This talk consists of work from the PhD thesis of Gabriel Martine La Boissoniere at McGill. Parts of the talk also involve joint work with S. Esedoglu (Michigan), K. Barmak (Columbia) and J.-P. Lessard (McGill).

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Subject: 

Recent Progress and Open Frontiers in Turing-Type Morphogenesis

Speaker: 
Andrew Krause
Date: 
Mon, May 10, 2021
Location: 
UBC
Online
Conference: 
PIMS Workshop on New Trends in Localized Patterns in PDES
Abstract: 

Motivated by recent work with biologists, I will showcase some mathematical results on Turing instabilities in complex domains. This is scientifically related to understanding developmental tuning in a variety of settings such as mouse whiskers, human fingerprints, bat teeth, and more generally pattern formation on multiple scales and evolving domains. Such phenomena are typically modelled using reaction-diffusion systems of morphogens, and one is often interested in emergent spatial and spatiotemporal patterns resulting from instabilities of a homogeneous equilibrium, which have been well-studied. In comparison to the well-known effects of how advection or manifold structure impacts unstable modes in such systems, I will present results on instabilities in heterogeneous systems, as well as those arising in the set-ting of evolving manifolds. These contexts require novel formulations of classical dispersion relations, and may have applications beyond developmental biology, such as in population dynamics (e.g. understanding colony or niche formation of populations in heterogeneous environments). These approaches also help close the vast gap between the simple theory of diffusion-driven pattern formation, and the messy reality of biological development, though there is still much work to be done in validating even complex theories against the rich dynamics observed in nature. I will close by discussing a range of open questions, many of which fall well beyond the extensions I will discuss.

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Subject: 

Localised patterns and semi-strong interaction, a unifying framework for reaction-diffusion systems

Speaker: 
Alan Champneys
Date: 
Mon, May 10, 2021
Location: 
UBC
Online
Conference: 
PIMS Workshop on New Trends in Localized Patterns in PDES
Abstract: 

Systems of activator-inhibitor reaction-diffusion equations posed on an infinite line are studied using a variety of analytical and numerical methods. A canonical form is considered that contains all known models with simple cubic autocatalytic nonlinearity and arbitrary constant and linear kinetics. Restricting attention at first to models that have a unique homogeneous equilibrium, this class includes the classical Schnakenberg and Brusselator models, as well as other systems proposed in the literature to model morphogenesis. Such models are known to feature Turing instability, when activator diffuses more slowly than inhibitor, leading to stable spatially periodic patterns. Conversely in the limit of small feed rates, semi-strong interaction asymptotic analysis as introduced by Michael Ward and his collaborators shows existence of isolated spike-like patterns.

Connecting these two regions, a certain universal two-parameter state diagram is revealed in which the Turing bifurcation becomes sub-critical, leading to the onset of homoclinic snaking. This regime then morphs into the spike regime, with the outer-fold being predicted by the semi-strong asymptotics. A rescaling of parameters and field concentrations shows how this state diagram can be studied independently of the diffusion rates. Temporal dynamics is found to strongly depend on the diffusion ratio though. A Hopf bifurcation occurs along the branch of stable spikes, which is subcritical for small diffusion ratio, leading to collapse to the homogeneous state. As the diffusion ratio increases, this bifurcation typically becomes supercritical, interacts with the homoclinic snaking and also with a supercritical homogeneous Hopf bifurcation, leading to complex spatio-temporal dynamics. The details are worked out for a number of different models that fit the theory using a mixture of weakly nonlinear analysis, semi-strong asymptotics and different numerical continuation algorithms.

The theory is extended include models, such as Gray-Scott, with bistability of homogeneous equilibria. A homotopy is studied that takes a Schnakenberg-like glycolysis model for r = 0 to the Gray-Scott model for r = 1. Numerical continuation is used to understand the complete sequence of transitions to two-parameter bifurcation diagrams within the localised pattern parameter regime as r varies. Several distinct codimension-two bifurcations are dis-covered including cusp and quadruple zero points for homogeneous steady states, a degenerate heteroclinic connection and a change in connectedness of the homoclinic snaking structure. The analysis is repeated for the Gierer-Meinhardt system, which lies outside the canonical framework. Similar transitions are found under homotopy between bifurcation diagrams for the case where there is a constant feed in the active field, to it being in the inactive field. Wider implications of the results are discussed for other kinds of pattern-formation systems as well as to distinguishing between different kinds of observed behaviour in the natural world.

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Subject: 

Localized slow patterns in singularly perturbed 2-component reaction-diffusion equations

Speaker: 
Arjen Doelman
Date: 
Mon, May 10, 2021
Location: 
UBC
Online
Conference: 
PIMS Workshop on New Trends in Localized Patterns in PDES
Abstract: 

Localized patterns in singularly perturbed reaction-diffusion equations typically consist of slow parts – in which the associated solution follows an orbit on a slow manifold in a reduced spatial dynamical system – alternated by fast excursions – in which the solution jumps from one slow manifold to another, or back to the original slow manifold. In this talk we consider the existence and stability of localized slow patterns that do not exhibit such jumps, i.e. that are completely embedded in a slow manifold of the singularly perturbed spatial dynamical system. These patterns have rarely been considered in the literature, for two reasons: (i) in the classical Gray-Scott/Gierer-Meinhardt type models that dominate the literature, the flow on the slow manifold is linear and thus cannot exhibit homoclinic pulse or heteroclinic front solutions; (ii) the slow manifolds occurring in the literature are typically trivial, or ‘vertical’ – i.e. given by u ≡ u_0, where u is the fast variable – so that the stability problem is determined by a simple (decoupled) scalar equation. The present talk is motivated by several explicit ecosystem models (of singularly perturbed reaction-diffusion type) that do give rise to nontrivial normally hyperbolic slow manifolds on which the flow may exhibit both homoclinic and heteroclinic orbits – that correspond to either stationary or traveling localized patterns. The associated spectral stability problems are at leading order given by a nonlinear, but scalar, eigenvalue problem with Sturm-Liouville type characteristics and we establish that homoclinic pulse patterns are typically unstable, while heteroclinic fronts can either be stable or unstable. However, we also show that homoclinic pulse patterns that are asymptotically close to a heteroclinic cycle may be stable. This result is obtained by explicitly determining the leading order approximations of 4 critical asymptotically small eigenvalues. Through this somewhat subtle analysis – that involves several orders of magnitude in the small parameter – we also obtain full control over the nature of the bifurcations – saddle-node, Hopf, global, etc. – that determine the existence and stability of the heteroclinic fronts and/or homoclinic pulses. Finally, we show that heteroclinic orbits may correspond to stable (slow) interfaces (in 2-dimensional space), while the homoclinic pulses must be unstable as localized stripes –even when they are stable in 1 space dimension.

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Subject: 

The Manhattan Curve and Rough Similarity Rigidity

Speaker: 
Ryokichi Tanaka
Date: 
Thu, May 20, 2021
Location: 
Zoom
Online
Conference: 
Pacific Dynamics Seminar
Abstract: 

For every non-elementary hyperbolic group, we consider the Manhattan curve, which was originally introduced by M. Burger (1993), associated to any pair of (say) word metrics. It is convex; we show that it is continuously differentiable and moreover is a straight line if and only if the corresponding two metrics are roughly similar, that is, they are within bounded distance after multiplying by a positive constant.

I would like to explain how it is related to central limit theorem for uniform counting measures on spheres, to ergodic theory of topological flows built on general hyperbolic groups, and to multifractal structure of Patterson-Sullivan measures. Furthermore I will present some explicit examples including a hyperbolic triangle group and compute the exact value of the mean distortion for a pair of word metrics by using automatic structures of the group.

Joint work with Stephen Cantrell (University of Chicago).

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Subject: 

Spike patterns as a window into non-injective transient diffusive processes

Speaker: 
Yana Nec
Date: 
Wed, May 12, 2021
Location: 
UBC
Online
Conference: 
PIMS Workshop on New Trends in Localized Patterns in PDES
Abstract: 

Complex natural systems at times manifest transitions between disparate diffusive regimes. Efforts to devise measurement techniques capable of identifying the cross-over moments have recently borne fruit, however interpretation of findings remains contentious when the bigger picture is considered. This study generalises the 1D Gierer-Meinhardt reaction – diffusion model to a system that permits transitions between regular diffusive regimes with distinct diffusivities as well as sub-diffusion of a variable order. This is a sufficiently general, yet tractable description for the dynamics of a pattern qualitatively redolent of molecular clusters subject to transient anomalous diffusion mechanisms. The resulting system of equations substantiates the difficulties encountered when attempting to distinguish between various diffusive regimes in experimental settings: a non-monotonic dependence of the pat-tern’s evolution on parameters defining the diffusion mechanism is a common occurrence, as is a non-injective mapping between a given sequence of diffusion regimes and ensuing drift behaviour.

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